In a game with two factions, theres no real villain i think. I think the main villain of the Sacred Order is Inngrim, while the main villain for Domination is King Venuin.
Maybe was a good idea that each race has his own villain of the opposed race, like, for humans, Nepherimas is the villain, because she is the queen of undead. For the Minotaurs, the villain is Chieftain Ursan.
For example:
Humans don't like undead and vice-versa;
Elfs don't like Infernal Demons and vice-versa;
Bearans don't like Minotaurs and vice-versa.
(And i don't know what to do about orcs and dwarfs lol)
That way, maybe each race could have an begginer area that your objective is try to kill the leader of the opposing race. Obviously you aren't going to kill the leader because thats would be too easy.